The Spider Cytogenetic Database

Current version: 12.5 (Feb 29, 2024)

Douglas Araujo, Marielle Cristina Schneider, Emygdio Paula-Neto and Doralice Maria Cella​
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Lycosidae  Sundevall, 1833

Cited as = species name adopted by the researcher that performed the cytogenetic analysis; 2n = diploid number of males and in parenthesis, diploid number of females; n = haploid number; 3n = triploid number; 4n = tetraploid number; SCS = sex chromosome system; T = telocentric; A = acrocentric; St = subtelocentric; Sm = submetacentric; M = metacentric; H = holocentric; S = supernumerary chromosome; (?) = dubious description; (*) = designation of the species name not recorded in the taxonomic database.

Valid name Cited as 2n SCS Chromosomal morphology Collection site Bibliography
Alopecosa aculeata (Clerck, 1757) Tarentula aculeata 28 X1X2 26A+X1X2A Finland Hackman, 1948
Alopecosa albofasciata (Brullé, 1832) ---- 28 X1X2 26A+X1X2A Israel Gorlova et al., 1997
Alopecosa pulverulenta (Clerck, 1757) Tarentula pulverulenta 28 X1X2 26A+X1X2A Finland Hackman, 1948
Anomalomma harishi Dyal, 1935 ---- 28 X1X2 ---- India Mittal, 1961
Anomalomma harishi Dyal, 1935 ---- 28 X1X2 26A+X1X2A India Mittal, 1963
Anomalomma sp. ---- 28 X1X2 26A+X1X2A India Sharma et al., 1958
Arctosa alpigena (Doleschall, 1852) ---- 26 X1X2 24A+X1X2A Finland Hackman, 1948
Arctosa alpigena (Doleschall, 1852) ---- (30) ---- 28A+2St Austria Dolejš et al., 2011
Arctosa alpigena lamperti Dhal, 1908 ---- 28 X1X2 24A+2St+X1X2A Czech Republic Dolejš et al., 2011
Arctosa cinerea (Fabricius, 1777) ---- 28 X1X2 26A+X1X2A Czech Republic Dolejš et al., 2011
Arctosa cinerea (Fabricius, 1777) ---- 28 X1X2 26A+X1X2 Turkey Kumbiçak et al., 2011
Arctosa figurata (Simon, 1876) ---- 28 X1X2 26A+X1X2A Czech Republic Dolejš et al., 2011
Arctosa leopardus (Sundevall, 1833) ---- 26 X1X2 24A+X1X2A Finland Hackman, 1948
Arctosa leopardus (Sundevall, 1833) ---- 28 X1X2 26A+X1X2A Czech Republic Dolejš et al., 2011
Arctosa lutetiana (Simon, 1876) Tricca lutetiana 28 X1X2 26A+X1X2A Czech Republic Dolejš et al., 2011
Arctosa maculata (Hahn, 1822) ---- 28 X`1X2 26A+X1X2A Czech Republic Dolejš et al., 2011
Arctosa mulani (Dyal, 1935) Pardosa mulani 28 X1X2 26A+X1X2A India Sharma et al., 1958
Arctosa perita (Latreille, 1799) ---- (12) ---- ---- Turkey Akan et al., 2005
Arctosa perita (Latreille, 1799) ---- 28 X1X2 26A+X1X2A Czech Republic Dolejš et al., 2011
Arctosa renidescens Buchar & Thaler, 1995 ---- 28 X1X2 26A+X1X2A Austria Dolejš et al., 2011
Arctosa sp. ---- 28 X1X2 ---- India Mittal, 1960
Arctosa sp. ---- 28 X1X2 26A+X1X2A India Mittal, 1963
Crocodilosa leucostigma (Simon, 1885) Lycosa leucostigma 28 X1X2 ---- India Srivastava & Shukla, 1986
Draposa lyrivulva (Bösenberg & Strand, 1906) Pardosa leucopalpis 28 X1X2 26A+X1X2A India Bole-Gowda, 1958
Draposa lyrivulva (Bösenberg & Strand, 1906) Pardosa leucopalpis 28 X1X2 26A+X1X2A India Mittal, 1963
Draposa lyrivulva (Bösenberg & Strand, 1906) Pardosa leucopalpis 24 X1X2 ---- India Srivastava & Shukla, 1986
Draposa oakleyi (Gravely, 1924) Pardosa lahorensis 28 X1X2 26A+X1X2A India Sharma et al., 1958
Draposa oakleyi (Gravely, 1924) Pardosa oakleyi 26 X1X2 ---- India Srivastava & Shukla, 1986
Evippa praelongipes (O.Pickard-Cambridge, 1871) ---- 26 X1X2 24A+X1X2A India Sharma et al., 1958
Evippa praelongipes (O.Pickard-Cambridge, 1871) ---- 26 X1X2 23A+1M+X1X2A Israel Gorlova et al., 1997
Gladicosa pulchra (Keyserling, 1877) Lycosa insopita 26+2S X1X2 ---- USA Montgomery, 1905
Hippasa agelenoides (Simon, 1884) ---- 28 X1X2 26A+X1X2A India Bole-Gowda, 1958
Hippasa agelenoides (Simon, 1884) ---- 24 X1X2 22A+X1X2A India Sharma et al., 1958
Hippasa agelenoides (Simon, 1884) ---- 28 X1X2 26A+X1X2A India Sharma & Ramakrishna, 2019
Hippasa deserticola Simon, 1889 Hippasa pisaurina 26 X1X2 ---- India Mittal, 1960
Hippasa deserticola Simon, 1889 Hippasa pisaurina 26 X1X2 24A+X1X2A India Mittal, 1963
Hippasa deserticola Simon, 1889 Hippasa madhuae 28 X1X2 26A+X1X2A India Parida et al., 1986
Hippasa deserticola Simon, 1889 Hippasa pisaurina 28 X1X2 ---- India Srivastava & Shukla, 1986
Hippasa holmerae Thorell, 1895 ---- 26 ---- ---- ---- Bugayong et al., 1999
Hippasa olivacea (Thorell, 1887) Hippasa oliracea 28 X1X2 26A+X1X2A India Parida & Sharma, 1987b
Hippasa olivacea (Thorell, 1887) ---- 28 X1X2 ---- India Parida & Sharma, 1987a
Hippasa olivacea (Thorell, 1887) ---- 28 X1X2 ---- India Sharma & Parida, 1987
Hippasa sp. ---- 22 X1X2 ---- India Parida & Sharma, 1987a
Hippasa sp. ---- 22 X1X2 ---- India Sharma & Parida, 1987
Hippasosa kumari (Dyal, 1935) Ocyale kumari 22 X1X2 20A+X1X2A India Sharma et al., 1958
Hogna sternalis (Bertkau, 1880) ---- 19 X 18T+XT Brazil Araujo et al., 2015a
Hygrolycosa rubrofasciata (Ohlert, 1865) ---- 20 X1X2 18A+X1X2A Russia Gorlov et al., 1995
Lycosa barnesi Gravely, 1924 ---- 27 X ---- India Srivastava & Shukla, 1986
Lycosa bistriata Gravely, 1924 ---- 28 X1X2 26A+X1X2A India Bole-Gowda, 1958
Lycosa carmichaeli Gravely, 1924 ---- 28 X1X2 ---- India Mittal, 1961
Lycosa carmichaeli Gravely, 1924 ---- 28 X1X2 26A+X1X2A India Mittal, 1963
Lycosa carmichaeli Gravely, 1924 ---- 22 X1X2 ---- India Srivastava & Shukla, 1986
Lycosa chaperi Simon, 1885 ---- 22 X1X2 20A+X1X2A India Mittal, 1963
Lycosa coelestis L. Koch, 1878 ---- 26 X1X2 24A+X1X2A Japan Suzuki, 1954
Lycosa erythrognatha Lucas, 1836 Lycosa erythrognata 22 X1X2 20A+X1X2A ---- Diaz & Saez, 1966
Lycosa erythrognatha Lucas, 1836 ---- 22 X1X2 20T+X1X2T Argentina Chemisquy et al., 2008
Lycosa erythrognatha Lucas, 1836 ---- 22 X1X2 20T+X1X2T Brazil Araujo et al., 2015a
Lycosa erythrognatha Lucas, 1836 ---- 22 X1X2 20A+X1X2A Brazil Cavenagh et al., 2022
Lycosa madani Pocock, 1901 ---- 24 X1X2 22A+X1X2A India Mittal, 1963
Lycosa nigrotibialis Simon, 1884 ---- 28 X1X2 ---- India Mittal, 1961
Lycosa nigrotibialis Simon, 1884 ---- 28 X1X2 26A+X1X2A India Mittal, 1963
Lycosa nigrotibialis Simon, 1884 ---- 24 X1X2 ---- India Srivastava & Shukla, 1986
Lycosa nordenskjoldi Tullgren, 1905 Lycosa nordenskölii 19 X 18A+XA ---- Diaz & Saez, 1966
Lycosa nordenskjoldi Tullgren, 1905 ---- 19 X 18T+XT Brazil Araujo et al., 2015a
Lycosa gr. nordenskjoldi ---- 19 X 18A+XA Brazil Cavenagh et al., 2022
Lycosa pampeana Holmberg, 1876 ---- 22 X1X2 20T+X1X2T Argentina Chemisquy et al., 2008
Lycosa phipsoni Pocock, 1899 ---- ---- X1X2 ---- India Mittal, 1961
Lycosa praegrandis C.L. Koch, 1836 Lycosa cf. nordmanni 22 X1X2 20A+X1X2A Israel Gorlova et al., 1997
Lycosa sericovittata Mello-Leitão, 1939 ---- 22 X1X2 20T+X1X2T Brazil Araujo et al., 2015a
Lycosa sericovittata Mello-Leitão, 1939 ---- (26) ---- 26T Brazil Araujo et al., 2015a
Lycosa sericovittata Mello-Leitão, 1939 ---- 22 X1X2 20A+X1X2A Brazil Cavenagh et al., 2022
Lycosa tarantula (Linnaeus, 1758) Lycosa narborensis 18 ---- ---- Turkey Akan et al., 2005
Lycosa sp. ---- 20-24? ---- ---- France Carnoy, 1885
Lycosa sp. ---- ---- X ---- ---- Chickering & Hard, 1935
Lycosa sp. ---- 28 X1X2 26A+X1X2A Japan Suzuki, 1954
Lycosa sp. ---- 28 X1X2 26A+X1X2A India Bole-Gowda, 1958
Lycosa sp. ---- 28 X1X2 26A+X1X2A India Sharma et al., 1958
Lycosa sp. ---- 28 X1X2 ---- ---- Sokolov, 1960
Lycosa sp. ---- 28 X1X2 26A+X1X2A India Mittal, 1963
Lycosa sp. ---- 22 X1X2 20T+X1X2T Uruguay Brum-Zorrilla & Postiglioni, 1980
Lycosa sp. (grupo thorelli) ---- 23 X1X2X3 20T+X1X2X3T Uruguay Postiglioni & Brum-Zorrilla, 1981
Lycosa sp. ---- 22 X1X2 20T+X1X2T Uruguay Postiglioni & Brum-Zorrilla, 1981
Lycosa sp. ---- 18 X1X2 ---- India Srivastava & Shukla, 1986
Lycosa sp. ---- 28 X1X2 ---- India Srivastava & Shukla, 1986
Lycosa sp. ---- 22 X1X2 20A+X1X2A India Parida & Sharma, 1987b
Lycosa sp. ---- 22 X1X2 ---- India Parida & Sharma, 1987a
Lycosa sp. ---- 28 X1X2 ---- India Parida & Sharma, 1987a
Lycosa sp. ---- 22 X1X2 ---- India Sharma & Parida, 1987
Lycosa sp. ---- 28 X1X2 ---- India Sharma & Parida, 1987
Lycosa sp. ---- 21 X1X2Y 16T+2St+X1X2M/Sm+YT Peru Navia et al., 2006
Margonia himalayensis (Gravely, 1924) Venonia himalayensis 28 X1X2 ---- India Mittal, 1961
Margonia himalayensis (Gravely, 1924) Venonia himalayensis 28 X1X2 26A+X1X2A India Mittal, 1963
Pardosa agrestis (Westring, 1861) ---- 28 X1X2 26A+X1X2A Russia Gorlov et al., 1995
Pardosa agricola (Thorell, 1856) Lycosa fluviatilis 28 X1X2 26A+X1X2A Finland Hackman, 1948
Pardosa alacris (C.L. Koch, 1833) ---- 28 X1X2 26A+X1X2A Czech Republic Kumbiçak et al., 2009
Pardosa amentata (Clerck, 1757) Lycosa saccata 28 X1X2 26A+X1X2A Finland Hackman, 1948
Pardosa amentata (Clerck, 1757) Lycosa paludicola 28 X1X2 26A+X1X2A Finland Hackman, 1948
Pardosa amentata (Clerck, 1757) Lycosa pullata 28 X1X2 26A+X1X2A Finland Hackman, 1948
Pardosa amentata (Clerck, 1757) ---- 28 X1X2 ---- ---- Sokolov, 1960
Pardosa astrigera L. Koch, 1878 Lycosa T-insignita 28 X1X2 ---- ---- Suzuki, 1950a
Pardosa astrigera L. Koch, 1878 Lycosa T-insignita 28 X1X2 26A+X1X2A Japan Suzuki, 1954
Pardosa astrigera L. Koch, 1878 Lycosa cinereofusca 28 X1X2 26A+X1X2A Japan Suzuki, 1954
Pardosa astrigera L. Koch, 1878 Pardosa T-insignita 28 X1X2 ---- Japan Matsumoto, 1977
Pardosa astrigera L. Koch, 1878 ---- 28 X1X2 26T+X1X2T Japan Kageyama et al., 1978
Pardosa astrigera L. Koch, 1878 ---- 28 X1X2 26T+X1X2T China Zhenling et al., 1996
Pardosa basiri (Dyal, 1935) Lycosa basiri 22 X1X2 ---- India Mittal, 1960
Pardosa basiri (Dyal, 1935) Lycosa basiri 22 X1X2 20A+X1X2A India Mittal, 1963
Pardosa bifasciata (C.L. Koch, 1834) ---- 28 X1X2 26A+X1X2A Turkey Kumbiçak et al., 2011
Pardosa birmanica Simon, 1884 Lycosa birmanica 28 X1X2 26A+X1X2A India Bole-Gowda, 1958
Pardosa birmanica Simon, 1884 ---- 28 X1X2 ---- India Datta & Chatterjee, 1983
Pardosa birmanica Simon, 1884 Lycosa birmanica 28 X1X2 ---- India Srivastava & Shukla, 1986
Pardosa birmanica Simon, 1884 ---- 28 X1X2 26A+X1X2A India Parida & Sharma, 1987b
Pardosa birmanica Simon, 1884 ---- 28 X1X2 ---- India Parida & Sharma, 1987a
Pardosa birmanica Simon, 1884 ---- 28 X1X2 ---- India Sharma & Parida, 1987
Pardosa birmanica Simon, 1884 ---- 28 X1X2 26T+X1X2T India Datta & Chatterjee, 1989
Pardosa fletcheri (Gravely, 1924) Lycosa fletcheri 28 X1X2 ---- India Srivastava & Shukla, 1986
Pardosa laura Karsch, 1879 ---- 28 X1X2 26T+X1X2T Japan Kageyama et al., 1978
Pardosa lugubris (Walckenaer, 1802) ---- 28 X1X2 26A+X1X2A Russia Gorlov et al., 1995
Pardosa morosa (L. Koch, 1870) ---- 26 X1X2 24A+X1X2A Czech Republic/Slovakia Král et al., 2011
Pardosa morosa (L. Koch, 1870) ---- 25? X1X2? 24A+1M? Czech Republic Král et al., 2011
Pardosa palustris (Linnaeus, 1758) Lycosa monticola 28 X1X2 26A+X1X2A Finland Hackman, 1948
Pardosa palustris (Linnaeus, 1758) Lycosa tarsalis 28 X1X2 24A+2M+X1X2A Finland Hackman, 1948
Pardosa palustris (Linnaeus, 1758) ---- 28 X1X2 26A+X1X2A Russia Gorlov et al., 1995
Pardosa plumipes (Thorell, 1875) ---- 28 X1X2 26A+X1X2A Russia Gorlov et al., 1995
Pardosa pseudoannulata (Bösenberg & Strand, 1906) Lycosa pseudoannulatus 28 X1X2 26A+X1X2A Japan Suzuki, 1954
Pardosa pseudoannulata (Bösenberg & Strand, 1906) Lycosa annandalei 28 X1X2 26A+X1X2A India Bole-Gowda, 1958
Pardosa pseudoannulata (Bösenberg & Strand, 1906) Lycosa pseudoannulata 28 X1X2 26T+X1X2T Japan Kageyama et al., 1978
Pardosa pseudoannulata (Bösenberg & Strand, 1906) Lycosa annandalei 28 X1X2 ---- India Srivastava & Shukla, 1986
Pardosa saltans Töpfer-Hofmann, 2000 ---- 28 X1X2 26A+X1X2A Czech Republic Kumbiçak et al., 2009
Pardosa sumatrana (Thorell, 1890) Lycosa sumatrana 24 X1X2 22A+X1X2A India Sharma, 1961
Pardosa sumatrana (Thorell, 1890) Lycosa sumatrana 28 X1X2 ---- India Srivastava & Shukla, 1986
Pardosa wagleri (Hahn, 1822) ---- 28 X1X2 26A+X1X2A ---- Kořínková & Král, 2013
Pardosa sp. ---- ---- X ---- ---- Chickering & Hard, 1935
Pardosa sp. ---- 28 X1X2 26T+X1X2T India Sharma & Gupta, 1956
Pardosa sp. ---- 28 X1X2 26A+X1X2A India Bole-Gowda, 1958
Pardosa sp. ---- 28 X1X2 ---- India Mittal, 1960
Pardosa sp. ---- 28 X1X2 26A+X1X2A India Mittal, 1963
Pirata piraticus (Clerck, 1757) ---- 26 X1X2 24A+X1X2A Finland Hackman, 1948
Pirata piraticus (Clerck, 1757) ---- 26 X1X2 24T+X1X2T ---- Yu et al., 1998a
Pirata punctipes (Gravely, 1924) Lycosa punctipes 28 X1X2 26A+X1X2A India Sharma, 1961
Pirata subpiraticus (Bösenberg & Strand, 1906) ---- 26 X1X2 24T+X1X2T Japan Kageyama et al., 1978
Pirata subpiraticus (Bösenberg & Strand, 1906) ---- 26 X1X2 24T+X1X2T ---- Yu et al., 1998b
Pirata sp. ---- 26 X1X2 ---- ---- Sokolov, 1960
Piratula latitans (Blackwall, 1841) Pirata latitans 24 X1X2 22A+X1X2A India Mittal, 1963
Piratula piratoides (Bösenberg & Strand, 1906) Pirata piratoides 26 X1X2 24T+X1X2T ---- Zhenling et al., 1997a
Piratula procurva (Bösenberg & Strand, 1906) Pirata procurvus 26 X1X2 24T+X1X2T Japan Kageyama et al., 1978
Piratula uliginosa (Thorell, 1856) Pirata uliginosus 24 X1X2 22A+X1X2A Finland Hackman, 1948
Rabidosa rabida (Walckenaer, 1837) Lycosa rabida 28 X1X2 26T+X1X2T USA Wise, 1983
Rabidosa rabida (Walckenaer, 1837) Lycosa rabida 28 X1X2 ---- USA Tugmon et al., 1990
Rabidosa rabida (Walckenaer, 1837) Lycosa rabida 28 X1X2 26T+X1X2T USA Wise & Taylor, 1995
Schizocosa bilineata (Emerton, 1885) ---- ---- X ---- ---- Chickering & Hard, 1935
Schizocosa communis (Emerton, 1885) Lycosa communis 22 X1X2 ---- ---- Painter, 1914
Schizocosa crassipes (Walckenaer, 1837) ---- 22 X1X2 ---- USA Hard, 1939
Schizocosa malitiosa (Tullgren, 1905) Lycosa malitiosa 22 X1X2 20T+X1X2T Uruguay Brum-Zorrilla & Cazenave, 1974
Schizocosa malitiosa (Tullgren, 1905) Lycosa malitiosa 22 X1X2 ---- Uruguay Benavente & Wettstein, 1977
Schizocosa malitiosa (Tullgren, 1905) Lycosa malitiosa 22 X1X2 20T+X1X2T Uruguay Brum-Zorrilla & Postiglioni, 1980
Schizocosa malitiosa (Tullgren, 1905) ---- 22 X1X2 20T+X1X2T Argentina Chemisquy et al., 2008
Schizocosa ocreata (Hentz, 1844) ---- 22 X1X2 ---- USA Stratton, 1997
Schizocosa rovneri Uetz & Dondale, 1979 ---- 22 X1X2 ---- USA Stratton, 1997
Schizocosa stridulans Stratton, 1984 ---- 22 X1X2 ---- USA Stratton, 1997
Schizocosa sp. ---- 28 X1X2 ---- India Mittal, 1960
Schizocosa sp. ---- 28 X1X2 26A+X1X2A India Mittal, 1963
Schizocosa sp. (grupo malitiosa) Lycosa sp. (grupo malitiosa) 23 X 22T+XM Uruguay Postiglioni & Brum-Zorrilla, 1981
Serratacosa himalayensis (Gravely, 1924) Lycosa himalayensis 28 X1X2 26A+X1X2A India Mittal, 1963
Tigrosa georgicola (Walckenaer, 1837) Lycosa georgicola 28 X1X2 24T+2A+X1X2T USA Wise, 1983
Tigrosa helluo (Walckenaer, 1837) Lycosa helluo 28 X1X2 26T+X1X2T USA Wise & Taylor, 1995
Trochosa ruricola (De Geer, 1778) ---- 26 X1X2 24A+X1X2A Finland Hackman, 1948
Trochosa ruricola (De Geer, 1778) Trachosa ruricula 20 X1X2 ---- Turkey Taşdemir et al., 2012
Trochosa spinipalpis (F.O.Pickard-Cambridge, 1895) ---- 26 X1X2 24A+X1X2A Finland Hackman, 1948
Trochosa spinipalpis (F.O.Pickard-Cambridge, 1895) ---- 26 X1X2 24A-T+X1X2A-T ---- Zhenling et al., 1997b
Tropicosa thorelli (Keyserling, 1877) Lycosa thorelli 22 X1X2 20T+X1X2T Uruguay Brum-Zorrilla & Postiglioni, 1980
Tropicosa thorelli (Keyserling, 1877) Lycosa thorelli 22 X1X2 20T+X1X2T Uruguay Postiglioni & Brum-Zorrilla, 1981
Venonia sp. ---- 26 X1X2 ---- India Mittal, 1961
Venonia sp. ---- 26 X1X2 ---- India Mittal, 1961
Venonia sp. ---- 26 X1X2 24A+X1X2A India Mittal, 1963
Wadicosa fidelis (O.Pickard-Cambridge, 1872) ---- 28 X1X2 26A+X1X2A Canary Island, Israel Forman et al., 2013
Wadicosa quadrifera (Gravely, 1924) Lycosa quadrifer 27 X ---- India Srivastava & Shukla, 1986
Xerolycosa miniata (C.L. Koch, 1834) ---- 22 X1X2 20A+X1X2A Finland Hackman, 1948
Xerolycosa miniata (C.L. Koch, 1834) ---- 22 X1X2 20A+X1X2A Russia Gorlov et al., 1995
Xerolycosa miniata (C.L. Koch, 1834) ---- 22 X1X2 20A+X1X2A Czech Republic Dolejš et al., 2011
Xerolycosa nemoralis (Westring, 1861) ---- 26 X1X2 24A+X1X2A Finland Hackman, 1948
Xerolycosa nemoralis (Westring, 1861) ---- 22 X1X2 20A+X1X2A Russia Gorlov et al., 1995
Xerolycosa nemoralis (Westring, 1861) ---- 22 X1X2 20A+X1X2A Czech Republic Dolejš et al., 2011