The Scorpion Cytogenetic Database

Current version: 13 (October 20, 2025)

Marielle Cristina Schneider, Viviane Fagundes Mattos and Doralice Maria Cella
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See all families
Bibliography
Buthidae   C.L. Koch, 1837 

Cited as = species name adopted by the researcher that performed the cytogenetic analysis; 2n = diploid number of males and in parenthesis, diploid number of females; M = metacentric; SM = submetacentric; ST = subtelocentric; A = acrocentric; T = telocentric; ? = dubious description.

Valid name Cited as 2n Chromosomal morphology Collection site Bibliography
Aegeobuthus gibbosus (Brullé, 1832) Mesobuthus gibbosus 28 ---- Turkey Karataş et al., 2019
Ananteris balzanii (Thorell, 1891) ---- 12 ---- Brazil Piza, 1947a
A. balzanii (Thorell, 1891) ---- 14 Holocentric Brazil Mattos et al., 2013
Androctonus amoreuxi (Audouin, 1826) ---- 24 Holocentric Egypt Moustafa et al., 2005
Androctonus amoreuxi (Audouin, 1826) ---- 24 Holocentric Morocco Šťáhlavský et al., 2020
Androctonus australis (Linnaeus, 1758) ---- 24 Holocentric Egypt Moustafa et al., 2005
Androctonus argoubi Kovařík, Lowe & Šťáhlavský, 2025 ---- 24 Holocentric Morocco Kovařík et al., 2025c
A. australis (Linnaeus, 1758) ---- 24 Holocentric Egypt Sadílek et al., 2015
Androctonus bicolor (Ehrenberg, 1828) ---- 24 Holocentric Egypt Moustafa et al., 2005
Androctonus bicolor (Ehrenberg, 1828) ---- 24 Holocentric Israel Šťáhlavský et al., 2020
Androctonus bourdoni Vachon, 1948 ---- 24 Holocentric Morocco Sadílek et al., 2015
Androctonus bourdoni Vachon, 1948 ---- 24 Holocentric Morocco Šťáhlavský et al., 2020
Androctonus crassicauda (Olivier, 1807) ---- 24 Holocentric Egypt Moustafa et al., 2005
A. crassicauda (Olivier, 1807) ---- 24 Holocentric Turkey Sadílek et al., 2015
Androctonus gonneti Vachon, 1948 ---- 24 Holocentric Morocco Šťáhlavský et al., 2020
Androctonus liouvillei (Pallary, 1924) ---- 24 Holocentric Morocco Šťáhlavský et al., 2020
Androctonus maelfaiti Lourenço, 2005 ---- 24 Holocentric Pakistan Sadílek et al., 2015
Androctonus mauritanicus (Pocock, 1902) Androctonus mauretanicus 24 ---- ---- Chovet et al., 1971
A. mauritanicus (Pocock, 1902) Androctonus mauretanicus 24 ---- ---- Moustafa et al., 2005
A. mauritanicus (Pocock, 1902) ---- 24 Holocentric Morocco Šťáhlavský et al., 2020
Androctonus sergenti Vachon, 1948 ---- 24 Holocentric Morocco Šťáhlavský et al., 2020
Androctonus stockmanni Kovařík, Lowe & Šťáhlavský, 2025 ---- 24 Holocentric Morocco Kovařík et al., 2025c
Androctonus tenuissimus Teruel, Kovařík & Turiel, 2013 ---- 24 Holocentric Egypt Šťáhlavský et al., 2020
Androctonus vaneki Kovařík, Lowe & Šťáhlavský, 2025 ---- 24 Holocentric Morocco Kovařík et al., 2025c
Apistobuthus pterygocercus Finnegan, 1932 ---- 22 Holocentric Oman Šťáhlavský et al., 2020
Babycurus buettneri Karsch, 1886 ---- 28 Holocentric Cameroon Kovařík et al., 2018a
Babycurus gigas Kraepelin, 1896 ---- 30 Holocentric Tanzania Kovařík et al., 2018a
Babycurus jacksoni (Pocock, 1890) ---- 16 Holocentric Tanzania Kovařík et al., 2018a
Barbaracurus sofomarensis Kovařík, Lowe, Seiter, Plíšková, Šťáhlavský, 2015 Babycurus sofomarensis 22 Holocentric Ethiopia Kovařík et al., 2015a
Barbaracurus sofomarensis Kovařík, Lowe, Seiter, Plíšková, Šťáhlavský, 2015 Babycurus sofomarensis 22 Holocentric Ethiopia Kovařík et al., 2018a
Barbaracurus somalicus (Hirst, 1907) ---- 36 Holocentric Somaliland Kovařík et al., 2018a
Barbaracurus zambonelli (Borelli, 1902) ---- 26 Holocentric Eritrea Kovařík et al., 2018a
Barbaracurus yemenensis Kovařík et al., 2018 ---- 26 Holocentric Saudi Arabia Kovařík et al., 2022b
Barbaracus feti Kovařík et al., 2019 ---- 23 Holocentric Somaliland Kovařík et al., 2019a
Buthacus macrocentrus (Ehrenberg, 1828) ---- 28 ---- Turkey Koç et al., 2019
Buthacus macrocentrus (Ehrenberg, 1828) ---- 28 Holocentric Turkey Šťáhlavský et al., 2020
Buthacus nigroaculeatus Levy, Amitai & Shulov, 1973 ---- 26 Holocentric Oman Šťáhlavský et al., 2020
Buthacus stockmanni Kovařík, Lowe & Šťáhlavský, 2016 ---- 20 Holocentric Morocco Kovařík et al., 2016a
Buthacus stockmanni Kovařík, Lowe & Šťáhlavský, 2016 ---- 20 Holocentric Morocco Šťáhlavský et al., 2020
Buthoscorpio sarasinorum (Karsh, 1892) ---- 14 Holocentric Sri Lanka Kovařík et al., 2016b
Buthus atlantis Pocock, 1889 ---- 22 Holocentric Morroco Šťáhlavský et al., 2020
Buthus awashensis Kovařík, 2011 ---- 22 ---- Ethiopia Kovařík et al., 2020b
Buthus awashensis Kovařík, 2011 ---- 22 Holocentric Ethiopia Šťáhlavský et al., 2020
Buthus berberensis Pocock, 1900 ---- 21 ---- Somaliland Kovařík et al., 2020b
Buthus maroccanus Birula, 1903 ---- 22 Holocentric Morocco Šťáhlavský et al., 2020
Buthus occitanus (Amoreux, 1789) ---- 22 ---- France Guénin, 1961
B. occitanus (Amoreux, 1789) Scorpio occitanus 44 ---- ---- Carnoy, 1885
B. occitanus (Amoreux, 1789) Scorpio occitanus 56 ---- ---- Makino, 1956
Buthus pococki Kovařík, Šťáhlavský & Elmi, 2020 ---- 22 ---- Somaliland Kovařík et al., 2020b
Buthus pygmaeus Kovařík, Lowe, Šťáhlavský & Elmi, 2025 ---- 22 Holocentric Somaliland Kovařík et al., 2025a
Buthus zeylensis Pocock, 1900 ---- 22 ---- Somaliland Kovařík et al., 2020b
Buthus zeylensis Pocock, 1900 ---- 22 Holocentric Somaliland Šťáhlavský et al., 2020
Centruroides baracoae Armas, 1976 ---- 26 Holocentric Cuba Šťáhlavský et al., 2020
Centruroides exilicauda (Wood, 1863) Centrurus exilicauda 26 ---- ---- Wilson, 1931
Centruroides gracilis (Latreille, 1804) ---- 24 Holocentric Cuba Šťáhlavský et al., 2020
Centruroides vittatus (Say, 1821) ---- 26 Holocentric United States Riess et al., 1978
Charmus laneus Karsh, 1879 ---- 9 Holocentric Sri Lanka Kovařík et al., 2016b
Compsobuthus acutecarinatus Simon, 1882 ---- 22 ---- Oman Kovařík et al., 2020a
Compsobuthus acutecarinatus Simon, 1882 ---- 22 Holocentric Oman Šťáhlavský et al., 2020
Compsobuthus arabicus Levy, Amitai & Shulov, 1973 ---- 22 ---- Oman Kovařík et al., 2020a
Compsobuthus arabicus Levy, Amitai & Shulov, 1973 ---- 22 Holocentric Oman Šťáhlavský et al., 2020
Compsobuthus eritreaensis Kovarík, Lowe, Plísková et Stahlavský, 2016 ---- 22 Holocentric Eritrea Kovařík et al., 2016c
Compsobuthus eritreaensis Kovarík, Lowe, Plísková et Stahlavský, 2016 ---- 22 Holocentric Eritrea Šťáhlavský et al., 2020
Compsobuthus maindroni (Kraepelin, 1901) ---- 22 ---- Oman Kovařík et al., 2020a
Compsobuthus maindroni (Kraepelin, 1901) ---- 22 Holocentric Oman Šťáhlavský et al., 2020
Compsobuthus matthiesseni (Birula, 1905) ---- 22 Holocentric Turkey Šťáhlavský et al., 2014
Compsobuthus matthiesseni (Birula, 1905) ---- 22 Holocentric Turkey Šťáhlavský et al., 2020
Compsobuthus schmiedeknechti Vachon, 1949 ---- 22 Holocentric Turkey Šťáhlavský et al., 2020
Compsobuthus ullrichi Kovařík, Lowe, Stockmann & Šťáhlavský, 2020 ---- 22 ---- Egypt Kovařík et al., 2020a
Gint amoudensis Kovařík, Lowe, Just, Awale, Elmi & Šťáhlavský, 2018 ---- 35, 36 ---- Somaliland Kovařík et al., 2018c
G. amoudensis Kovařík, Lowe, Just, Awale, Elmi & Šťáhlavský, 2018 ---- 35, 36 Holocentric Somaliland Just et al., 2022
Gint banfasae Kovařík & Lowe, 2019 ---- 18, 19 Holocentric Somaliland Just et al., 2022
Gint dabakalo Kovařík & Mazuch, 2015 ---- 23 ---- Somaliland Kovařík et al., 2018c
G. dabakalo Kovařík & Mazuch, 2015 ---- 23, 24, 27 Holocentric Somaliland Just et al., 2022
Gint gaitako Kovarik, Lowe, Pliskova & Stahlavsky, 2013 ---- 30 Holocentric Ethiopia Kovařík et al., 2013
G. gaitako Kovarik, Lowe, Pliskova & Stahlavsky, 2013 ---- 30 Holocentric Ethiopia Just et al., 2022
Gint gubanensis Kovařík et al., 2018 ---- 45 Holocentric Somaliland Just et al., 2022
Gint maidensis Kovařík, Lowe, Just, Awale, Elmi & Šťáhlavský, 2018 ---- 34 ---- Somaliland Kovařík et al., 2018c
G. maidensis Kovařík, Lowe, Just, Awale, Elmi & Šťáhlavský, 2018 ---- 34 Holocentric Somaliland Just et al., 2022
Grosphus sp. ---- 13 Holocentric Madagascar Šťáhlavský et al., 2020
Hemilychas alexandrinus (Hirst, 1911) Lychas alexandrinus 14 Holocentric Australia Shanahan, 1989a
Heteroctenus abudi (Armas & Marcano Fondeur, 1987) ---- 16 Holocentric Dominican Republic Šťáhlavský et al., 2020
Heteroctenus garridoi (Armas, 1974) ---- 23 Holocentric Cuba Šťáhlavský et al., 2020
Heteroctenus princeps (Karsch, 1879) ---- 16 Holocentric Dominican Republic Šťáhlavský et al., 2020
Hottentotta arenaceus (Purcell, 1902) ---- 14 Holocentric Namibia Šťáhlavský et al., 2020
Hottentotta conspersus (Thorell, 1876) ---- 16 Holocentric Namibia Šťáhlavský et al., 2020
Hottentotta flavidulus Teruel & Rein, 2010 ---- 24 Holocentric Namibia Šťáhlavský et al., 2020
Hottentotta hottentotta (Fabricius, 1787) ---- 16 Holocentric Mali Šťáhlavský et al., 2020
Hottentotta judaicus (Simon, 1872) ---- 16 6M+10A West Bank Qumsiyeh et al., 2013
Hottentotta minusalta Vachon, 1959 ---- 16 Holocentric Afghanistan Šťáhlavský et al., 2020
Hottentotta saulcyi (Simon, 1828) ---- 14 ---- Turkey Koç et al., 2019
Hottentotta saulcyi (Simon, 1828) ---- 14 Holocentric Turkey Šťáhlavský et al., 2020
Hottentotta sousai Turiel, 2014 ---- 14 Holocentric Morocco Šťáhlavský et al., 2020
Hottentotta tamulus (Fabricius, 1798) Buthus tamulus 22 Holocentric India Sharma et al., 1959
H. tamulus Buthus tamulus 20-28 Holocentric India Venkatanarasimhaiah & Rajasekarasetty, 1964a
H. tamulus Buthus tamulus 24 Holocentric India Gupta & Sarker, 1965
Hottentotta trilineatus (Peters, 1861) Buthotus trilineatus 24 ---- Zimbabwe Newlands & Martindale, 1980
Hottentotta ugandensis Kovařík, 2013 ---- 22 Holocentric Uganda Šťáhlavský et al., 2020
Hottentotta sp. ---- 22 Holocentric Pakistan Šťáhlavský et al., 2020
Ischnotelson guanambiensis (Lenarducci et al., 2005) ---- 25 Holocentric Brazil Ubinski et al., 2018
Ischnotelson peruassu Esposito et al., 2017 ---- 26 Holocentric Brazil Ubinski et al., 2018
I. peruassu Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017 ---- 26 Holocentric Brazil Lima et al., 2023
Ischnotelson sp. ---- 26 Holocentric Brazil Ubinski et al., 2018
Isometroides vescus (Karsch, 1880) ---- 14 Holocentric Australia Shanahan, 1989a
Isometrus maculatus (DeGeer, 1778) ---- 12 Holocentric Brazil Piza, 1947b
I. maculatus (DeGeer, 1778) ---- 12 Holocentric Brazil Piza, 1955
I. maculatus (DeGeer, 1778) ---- 12 ---- Brazil Piza, 1950a
I. maculatus (DeGeer, 1778) ---- 14 Holocentric Australia Shanahan, 1989a
Isometrus melanodactylus (L. Koch, 1867) ---- 14 Holocentric Australia Shanahan, 1989a
Isometrus thwaitesi Pocock, 1987 ---- 8 Holocentric Sri Lanka Kovařík et al., 2016b
Isometrus thwaitesi Pocock, 1987 ---- 8 Holocentric Sri Lanka Šťáhlavský et al., 2020
Jaguajir agamemnon (Koch, 1839) Rhopalurus agamemnon 28 Holocentric Brazil Mattos et al., 2013
J. agamemnon (Koch, 1839) ---- 28 Holocentric Brazil Ubinski et al., 2018
Jaguajir pintoi (Mello-Leitão 1932) ---- 28 Holocentric Brazil Ubinski et al., 2018
Jaguajir rochae (Borelli, 1910) Rhopalurus rochai 28 ---- Brazil Piza, 1957
J. rochae (Borelli, 1910) Rhopalurus rochai 28 Holocentric Brazil Mattos et al., 2013
J. rochae (Borelli, 1910) ---- 28 Holocentric Brazil Ubinski et al., 2018
Leirus abdullahbayram Yagmur, Koc & Kunt, 2009 ---- 22 Holocentric Turkey Šťáhlavský et al., 2014
Leirus abdullahbayram Yagmur, Koc & Kunt, 2009 ---- 22 Holocentric Turkey Šťáhlavský et al., 2020
Leiurus hebraeus (Birula, 1908) ---- 22 Holocentric Jordan Šťáhlavský et al., 2020
Leiurus quinquestriatus (Ehrenberg, 1828) ---- 22, XY ---- West Bank Qumsiyeh et al., 2013
Leiurus sp. ---- 22 Holocentric Africa Šťáhlavský et al., 2020
Lychas krali Kovařík, 1995 ---- 16 Holocentric Thailand Šťáhlavský et al., 2020
Lychas marmoreus (C. L. Koch, 1844) ---- 12, 14, 15 Holocentric Australia Shanahan, 1989a
L. marmoreus (C. L. Koch, 1844) ---- 12, 14, 15 Holocentric Australia Shanahan & Hayman, 1990
Lychas mucronatus (Fabricius, 1798) ---- 10 Holocentric Thailand Šťáhlavský et al., 2020
Lychas obsti Kraepelin, 1913 ---- 10 Holocentric Ethiopia, Tanzania Šťáhlavský et al., 2020
Lychas scutilus C. L. Koch, 1845 ---- 14 Holocentric Thailand Šťáhlavský et al., 2020
Lychas srilankensis Lourenço, 1997 ---- 16 Holocentric Sri Lanka Kovařík et al., 2016b
Lychas variatus (Thorell, 1876) ---- 14, 16 Holocentric Australia Shanahan, 1989a
L. variatus (Thorell, 1876) ---- 14, 16 Holocentric Australia Shanahan & Hayman, 1990
Mesobuthus eupeus (C. L. Koch, 1839) Buthus eupeus 22 ---- ---- Makino, 1956
M. eupeus (C.L. Koch, 1838) ---- 20 ---- Turkey Karataş et al., 2019
M. eupeus (C.L. Koch, 1838) ---- 22 Holocentric Turkey Šťáhlavský et al., 2020
M. eupeus (C.L. Koch, 1838) ---- 22 ---- Georgia Kovařík et al., 2022
Mesobuthus haarlovi Vachon, 1958 ---- 22 ---- Afghanistan Kovařík et al., 2022
Mesobuthus macmahoni (Pocock, 1900) Buthus macmahoni 24 ---- India Sharma et al., 1959
M. macmahoni (Pocock, 1900) ---- 22 ---- Pakistan Kovařík et al., 2022
Mesobuthus martensii (Karsch, 1879) Buthus martensii 24 ---- ---- Makino, 1956
M. martensii (Karsch 1879) Buthus martensii 24 ---- China Sato, 1940
Mesobuthus mesopotamicus (Penther, 1912) ---- 22 ---- Turkey Kovařík et al., 2022
Mesobuthus rahsenae Kovařík et al., 2022 ---- 20 ---- Turkey Kovařík et al., 2022
Microbuthus gardneri Lowe, 2010 ---- 26 Holocentric Oman Lowe et al., 2018
Microbuthus kristensenorum Lowe, 2010 ---- 26 Holocentric Oman Lowe et al., 2018
Microbuthus satyrus Lowe, Kovarik, Stockmann and Stahlavsky 2018 ---- 26 Holocentric Oman Lowe et al., 2018
Microbuthus satyrus Lowe, Kovarik, Stockmann and Stahlavsky 2018 ---- 26 Holocentric Yemen Lowe et al., 2018
Odontobuthus doriae (Thorell, 1876) Buthus doriae 22 ---- India Sharma et al., 1959
Orthochirus afar Kovařík, Lowe & Šťáhlavský, 2016 ---- 24 Holocentric Somaliland Šťáhlavský et al., 2020
Orthochirus glabrifrons (Kraepelin, 1903) ---- 22 Holocentric Oman Šťáhlavský et al., 2020
Orthochirus innesi Simon, 1910 ---- 22 Holocentric Morocco Šťáhlavský et al., 2020
Parabuthus abyssinicus Pocock, 1901 ---- 16 Holocentric Ethiopia Kovařík et al., 2016d
P. abyssinicus Pocock, 1901 ---- 16 Holocentric Eritrea Kovařík et al., 2016d
P. abyssinicus Pocock, 1901 ---- 16 Holocentric Ethiopia Šťáhlavský et al., 2020
Parabuthus capensis (Ehrenberg, 1831) ---- 20 Holocentric South Africa Šťáhlavský et al., 2020
Parabuthus dorisae Kovařík, Lowe, Elmi & Šťáhlavský, 2024 ---- 16 Holocentric Somaliland Kovařík et al., 2024c
Parabuthus glabrimanus Prendini & Esposito, 2010 ---- 20 Holocentric Namibia Šťáhlavský et al., 2020
Parabuthus kabateki Kovařík, Lowe, Elmi & Šťáhlavský, 2019 ---- 16 ---- Somaliland Kovařík et al., 2019d
Parabuthus kajibu Kovařík, Lowe, Plíšková & Šťáhlavský, 2016 ---- 18 Holocentric Ethiopia Kovařík et al., 2016d
Parabuthus kajibu Kovařík, Lowe, Plíšková & Šťáhlavský, 2016 ---- 18 Holocentric Ethiopia Šťáhlavský et al., 2020
Parabuthus mossambicensis (Peters, 1861) ---- 36 ---- Zimbabwe Newlands & Martindale, 1980
Parabuthus mossambicensis (Peters, 1861) ---- 18 Holocentric Mozambique Šťáhlavský et al., 2020
Parabuthus pallidus Pocock, 1895 ---- 20 Holocentric Ethiopia Kovařík et al., 2016d
Parabuthus pallidus Pocock, 1895 ---- 20 Holocentric Ethiopia Šťáhlavský et al., 2020
Parabuthus planicauda (Pocock, 1889) ---- 18 Holocentric South Africa Šťáhlavský et al., 2020
Parabuthus puntlandus Kovařík, Šťáhlavský, Elmi & Štarha, 2025 ---- 16 Holocentric Somalia Kovařík et al., 2025b
Parabuthus quincyae Kovařík, Lowe, Elmi & Šťáhlavský, 2024 ---- 16 Holocentric Somaliland Kovařík et al., 2024c
Parabuthus raudus (Simon, 1888) ---- 18 ---- Zimbabwe Newlands & Martindale, 1980
Parabuthus raudus (Simon, 1888) ---- 20 Holocentric Namibia Šťáhlavský et al., 2020
Parabuthus robustus Kovařík, Lowe, Elmi & Šťáhlavský, 2019 ---- 16 ---- Somaliland Kovařík et al., 2019d
Parabuthus somalilandus Kovařík, Lowe, Elmi & Šťáhlavský, 2019 ---- 16 ---- Somaliland Kovařík et al., 2019d
Parabuthus transvaalicus (Purcell, 1899) ---- 20 ---- Zimbabwe Newlands & Martindale, 1980
Physoctonus debilis (Koch 1840) ---- 26 Holocentric Brazil Ubinski et al., 2018
Reddyanus basilicus (Karsh, 1879) ---- 15, 16 Holocentric Sri Lanka Kovařík et al., 2016b
Reddyanus basilicus (Karsh, 1879) ---- 15, 16 Holocentric Sri Lanka Šťáhlavský et al., 2020
Reddyanus ceylonensis Kovařík, Lowe, Ranawana, Hoferek, Jayarathne, Plíšková et Šťáhlavský, 2016 ---- 16 Holocentric Sri Lanka Kovařík et al., 2016b
Reddyanus ceylonensis Kovařík, Lowe, Ranawana, Hoferek, Jayarathne, Plíšková et Šťáhlavský, 2016 ---- 16 Holocentric Sri Lanka Šťáhlavský et al., 2020
Reddyanus furai Kovařík and Šťáhlavský, 2019 ---- 14 Holocentric Vietnam Kovařík and Šťáhlavský, 2019
Reddyanus justi Kovařík, Lowe and Šťáhlavský, 2020 ---- 12 Holocentric Laos Kovařík et al., 2020c
Reddyanus loebli (Vachon, 1982) ---- 17 Holocentric Sri Lanka Kovařík et al., 2016b
Reddyanus loebli (Vachon, 1982) ---- 17 Holocentric Sri Lanka Šťáhlavský et al., 2020
Reddyanus majkusi Kovařík and Šťáhlavský, 2019 ---- 16 Holocentric Malaysia Kovařík and Šťáhlavský, 2019
Reddyanus rolciki Kovařík and Šťáhlavský, 2019 ---- 14 Holocentric Vietnam Kovařík and Šťáhlavský, 2019
Reddyanus rolciki Kovařík and Šťáhlavský, 2019 ---- 14 Holocentric Vietnam Šťáhlavský et al., 2020
Reddyanus schwotti Kovařík and Šťáhlavský, 2019 ---- 11 Holocentric Thailand Kovařík and Šťáhlavský, 2019
Reddyanus schwotti Kovařík and Šťáhlavský, 2019 ---- 11 Holocentric Thailand Šťáhlavský et al., 2020
Rhopalurus laticauda Thorell 1876 ---- 22 Holocentric Brazil Ubinski et al., 2018
Somalicharmus whitmanae Kovařík, 1998 ---- 20 Holocentric Ethiopia Kovařík et al., 2016e
S. whitmanae Kovařík, 1998 ---- 21 Holocentric Ethiopia Kovařík et al., 2016e
Thaicharmus sp. ---- 16 Holocentric Vietnam Šťáhlavský et al., 2020
Tityopsis rolandoi Kovařík, Stockmann, Šťáhlavský & Yong, 2024 ---- 20 Holocentric Cuba Kovařík et al., 2024a
Tityopsis sheylae Teruel & Rodriguez-Cabrera, 2020 ---- 22 Holocentric Cuba Kovařík et al., 2024a
Tityus aba Candido, Lucas, de Souza, Diaz & Lira-da-Silva, 2005 ---- 18 Holocentric Brazil Lima et al., 2023
Tityus apiacas Lourenço, 2002 ---- 14 Holocentric Brazil Lima et al., 2023
Tityus bahiensis (Perty, 1833) ---- 9 Holocentric Brazil Piza, 1947c
T. bahiensis (Perty, 1833) ---- 6 Holocentric Brazil Piza, 1939a
T. bahiensis (Perty, 1833) ---- 6 Holocentric Brazil Piza, 1939b
T. bahiensis (Perty, 1833) ---- 6, 9, 18 Holocentric Brazil Piza, 1940
T. bahiensis (Perty, 1833) ---- 6 Holocentric Brazil Brieger & Graner, 1943
T. bahiensis (Perty, 1833) ---- 5 Holocentric Brazil Piza, 1944
T. bahiensis (Perty, 1833) ---- 6, 10 Holocentric Brazil Piza, 1943a
T. bahiensis (Perty, 1833) ---- 6 Holocentric Brazil Piza, 1943b
T. bahiensis (Perty, 1833) ---- 9 Holocentric Brazil Piza, 1947c
T. bahiensis (Perty, 1833) ---- 7, 9 Holocentric Brazil Piza, 1948c
T. bahiensis (Perty, 1833) ---- 9, 10 Holocentric Brazil Piza, 1948b
T. bahiensis (Perty, 1833) ---- 17, 18, 19 Holocentric Brazil Piza, 1949
T. bahiensis (Perty, 1833) ---- 9, 10 Holocentric Brazil Piza, 1950a
T. bahiensis (Perty, 1833) ---- 10 Holocentric Brazil Takahashi, 1976
T. bahiensis (Perty, 1833) ---- 9, 10, 12, 16 Holocentric Brazil Piza, 1950b
T. bahiensis (Perty, 1833) ---- 10 Holocentric Brazil Cunha & Pavan, 1954
T. bahiensis (Perty, 1833) ---- 5, 6, 9, 10 Holocentric Brazil Schneider et al., 2009b
T. bahiensis (Perty, 1833) ---- 6 Holocentric Brazil Mattos et al., 2013
T. bahiensis (Perty, 1833) ---- 12, 13, 14, 15 Holocentric Argentina Adilardi et al., 2020
T. bahiensis (Perty, 1833) ---- 10 Holocentric Brazil Lima et al., 2023
Tityus carrilloi Ojanguren-Affilastro, 2021 Tityus trivittatus (6) Holocentric Argentina Adilardi et al., 2014
Tityus clathratus C. L. Kock, 184 ---- 19, 20 Holocentric Brazil Mattos et al., 2018
Tityus confluens (Borelli, 1899) ---- 13 Holocentric Brazil Mattos et al., 2013
T. confluens (Borelli, 1899) ---- 5 Holocentric` Argentina Adilardi et al., 2016
T. confluens (Borelli, 1899) ---- 6 Holocentric Argentina Adilardi et al., 2016
Tityus costatus (Karsch, 1879) ---- (16) Holocentric Brazil Mattos et al., 2013
Tityus crassimanus (Thorell, 1876) ---- 29 Holocentric Dominican Republic Šťáhlavský et al., 2020
Tityus curupi Ojanguren-Affilastro et al., 2017 ---- 32 Holocentric Argentina Ojanguren-Affilastro et al., 2017
Tityus ecuadoriensis Kraepelin, 1896 ---- 24 Holocentric Ecuador Šťáhlavský et al., 2020
Tityus elii (Armas & Marcano Fondeur, 1992) ---- 22 Holocentric Dominican Republic Šťáhlavský et al., 2020
Tityus fasciolatus (Pessôa, 1935) ---- 14 Holocentric Brazil Mattos et al., 2013
Tityus fuhrmanni Kraepelin, 1914 ---- 22 Holocentric Colombia Šťáhlavský et al., 2020
Tityus gaffini Lourenço, 2000 ---- 22 Holocentric Colombia Šťáhlavský et al., 2020
Tityus magnimanus (Pocock, 1897) ---- 20 Holocentric ---- Kovařík et al., 2009
Tityus mana Lourenço, 2012 ---- 20 Holocentric French Guiana Šťáhlavský et al., 2020
Tityus maranhensis (Lourenço, de Jesus Junior & Limeira-de-Oliveira, 2006) ---- 20 Holocentric Brazil Mattos et al., 2013
T. maranhensis (Lourenço, de Jesus Junior & Limeira-de-Oliveira, 2006) ---- 20 Holocentric Brazil Almeida et al., 2022
Tityus martinpaechi (Lourenço, 2001) ---- 6 ---- Brazil Mattos et al., 2013
Tityus matogrossensis (Borelli, 1901) ---- 20 ---- Brazil Piza, 1947a
Tityus mattogrossensis (Borelli, 1901) ---- 19, 20 Holocentric Brazil Mattos et al., 2018
Tityus metuendus Pocock, 1897 ---- 15, 16 ---- Brazil Piza, 1952
T. metuendus Pocock, 1897 ---- 14 Holocentric Brazil Lima et al., 2023
Tityus neglectus (Mello-Leitão, 1932) ---- 26, 27 ---- Brazil Piza, 1950b
Tityus neibae Armas, 1999 ---- 32 Holocentric Dominican Republic Šťáhlavský et al., 2020
Tityus obscurus (Gervais, 1843) ---- 11, 12, 13, 14, 16 Holocentric Brazil Almeida et al., 2017
Tityus paraguayensis (Kraepelin, 1895) ---- 16 ---- Brazil Mattos et al., 2013
Tityus paraguayensis (Kraepelin, 1895) ---- 16, 17, 18 Holocentric Brazil Mattos et al., 2018
Tityus pusillus Pocock, 1893 ---- 19, 20 Holocentric Brazil Mattos et al., 2018
Tityus serrulatus (Lutz & Mello, 1922) ---- (12) ---- Brazil Piza, 1947a
T. serrulatus (Lutz & Mello, 1922) ---- 12 Holocentric Brazil Schneider & Cella, 2010
T. serrulatus (Lutz & Mello, 1922) ---- 12 Holocentric Brazil Lima et al., 2020
Tityus silvestris Pocock, 1897 ---- 16, 24 Holocentric Brazil Mattos et al., 2018
T. silvestris Pocock, 1897 ---- 24 Holocentric Brazil Almeida et al., 2019
T. silvestris Pocock, 1897 ---- 24 Holocentric Brazil Almeida et al., 2022
Tityus smithii Pocock, 1893 ---- 6 Holocentric Grenada Šťáhlavský et al., 2020
Tityus stigmurus (Thorell, 1876) ---- (16) ---- Brazil Piza, 1950b
T. stigmurus (Thorell, 1876) ---- (16) ---- Brazil Mattos et al., 2013
Tityus trivittatus (Kraepelin, 1898) ---- 14 ---- Brazil Piza, 1948a
Tityus uruguayensis Borelli 1901 ---- 31 Holocentric Argentina Ojanguren-Affilastro et al., 2017
Tityus ythieri (Lourenço, 2007) ---- 20 Holocentric ---- Kovařík et al., 2009
Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha 1997) ---- 22 Holocentric Brazil Ubinski et al., 2018
Troglorhopalurus translucidus Lourenço et al., 2004 ---- 20 Holocentric Brazil Ubinski et al., 2018
Uroplectes carinatus (Pocok, 1890) ---- 20, 48 ---- Zimbabwe Newlands & Martindale, 1980
Uroplectes chubbi (Hirst, 1911) ---- 20 ---- Zimbabwe Newlands & Martindale, 1980
Uroplectes ebogo Kovařík, Šťáhlavský & Govorov, 2024 ---- 24 Holocentric Cameroon Kovařík et al., 2024b
Uroplectes emiliae (Werner, 1916) ---- 28 Holocentric Tanzania Šťáhlavský et al., 2020
Uroplectes fischeri (Karsch, 1879) ---- 28 Holocentric Ethiopia Kovařík et al., 2016f
Uroplectes flavoviridis (Peters, 1861) ---- 26 ---- Zimbabwe Newlands & Martindale, 1980
Uroplectes formosus Pocock, 1890 ---- 16 Holocentric South Africa Šťáhlavský et al., 2020
Uroplectes olivaceus (Pocock, 1896) ---- 24 ---- Zimbabwe Newlands & Martindale, 1980
Uroplectes otjimbinguensis (Karsch, 1879) ---- 26 Holocentric Namibia Šťáhlavský et al., 2020
Uroplectes planimanus (Karsch, 1879) ---- 22 ---- Zimbabwe Newlands & Martindale, 1980
Uroplectes vittatus (Thorell, 1876) ---- 24 ---- Zimbabwe Newlands & Martindale, 1980
Zabius fuscus Thorell, 1893 ---- 18 Holocentric Argentina Adilardi et al., 2015