The spider cytogenetic database

Current version: 6.5 (Jan 31, 2018)

Douglas Araujo, Marielle Cristina Schneider, Emygdio Paula-Neto and Doralice Maria Cella


Introduction of Database Families - Cytogenetics of spiders Bibliography Arthropoda cytogenetics group Scorpion database Pseudoscorpion database Harvestmen database Chromosomal analyses

 

 

Lycosidae Sundevall, 1833

 

Cited as = species name adopted by the researcher that performed the cytogenetic analysis; 2n = diploid number of males and in parenthesis, diploid number of females; n = haploid number;

3n = triploid number; 4n = tetraploid number; SCS = sex chromosome system; T = telocentric; A = acrocentric; St = subtelocentric; Sm = submetacentric; M = metacentric; H = holocentric;

S = supernumerary chromosome; (?) = dubious description; (*) = designation of the species name not recorded in the taxonomic database.

Valid name

Cited as

2n

SCS

Chromosomal morphology

Collection site

Bibliography

Alopecosa aculeata (Clerck, 1757)

Tarentula aculeata

28

X1X2

26A+X1X2A

Finland

Hackman, 1948

Alopecosa albofasciata (Brullé, 1832)

----

28

X1X2

26A+X1X2A

Israel

Gorlova et al., 1997

Alopecosa pulverulenta (Clerck, 1757)

Tarentula pulverulenta

28

X1X2

26A+X1X2A

Finland

Hackman, 1948

Anomalomma harishi Dyal, 1935

----

28

X1X2

----

India

Mittal, 1961

A. harishi Dyal, 1935

----

28

X1X2

26A+X1X2A

India

Mittal, 1963

Anomalomma sp.

----

28

X1X2

26A+X1X2A

India

Sharma et al., 1958

Arctosa alpigena (Doleschall, 1852)

----

26

X1X2

24A+X1X2A

Finland

Hackman, 1948

A. alpigena (Doleschall, 1852)

----

(30)

----

28A+2St

Austria

Dolejš et al., 2011

Arctosa alpigena lamperti Dhal, 1908

----

28

X1X2

24A+2St+X1X2A

Czech Republic

Dolejš et al., 2011

Arctosa cinerea (Fabricius, 1777)

----

28

X1X2

26A+X1X2A

Czech Republic

Dolejš et al., 2011

A. cinerea (Fabricius, 1777) ---- 28 X1X2 26A+X1X2 Turkey Kumbiçak et al., 2011

Arctosa figurata (Simon, 1876)

----

28

X1X2

26A+X1X2A

Czech Republic

Dolejš et al., 2011

Arctosa leopardus (Sundevall, 1833)

----

26

X1X2

24A+X1X2A

Finland

Hackman, 1948

A. leopardus (Sundevall, 1833)

----

28

X1X2

26A+X1X2A

Czech Republic

Dolejš et al., 2011

Arctosa lutetiana (Simon, 1876)

Tricca lutetiana

28

X1X2

26A+X1X2A

Czech Republic

Dolejš et al., 2011

Arctosa maculata (Hahn, 1822)

----

28

X`1X2

26A+X1X2A

Czech Republic

Dolejš et al., 2011

Arctosa mulani (Dyal, 1935)

Pardosa mulani

28

X1X2

26A+X1X2A

India

Sharma et al., 1958

Arctosa perita (Latreille, 1799)

----

(12)

----

----

Turkey

Akan et al., 2005

A. perita (Latreille, 1799)

----

28

X1X2

26A+X1X2A

Czech Republic

Dolejš et al., 2011

Arctosa renidescens Buchar & Thaler, 1995

----

28

X1X2

26A+X1X2A

Austria

Dolejš et al., 2011

Arctosa sp.

----

28

X1X2

----

India

Mittal, 1960

Arctosa sp.

----

28

X1X2

26A+X1X2A

India

Mittal, 1963

Crocodilosa leucostigma (Simon, 1885)

Lycosa leucostigma

28

X1X2

----

India

Srivastava & Shukla, 1986

Draposa lyrivulva (Bösenberg & Strand, 1906)

Pardosa leucopalpis

28

X1X2

26A+X1X2A

India

Bole-Gowda, 1958

D. lyrivulva (Bösenberg & Strand, 1906)

Pardosa leucopalpis

28

X1X2

26A+X1X2A

India

Mittal, 1963

D. lyrivulva (Bösenberg & Strand, 1906)

Pardosa leucopalpis

24

X1X2

----

India

Srivastava & Shukla, 1986

Draposa oakleyi (Gravely, 1924)

Pardosa lahorensis

28

X1X2

26A+X1X2A

India

Sharma et al., 1958

D. oakleyi (Gravely, 1924)

Pardosa oakleyi

26

X1X2

----

India

Srivastava & Shukla, 1986

Evippa praelongipes (O.Pickard-Cambridge, 1871)

----

26

X1X2

24A+X1X2A

India

Sharma et al., 1958

E. praelongipes (O.Pickard-Cambridge, 1871)

----

26

X1X2

23A+1M+X1X2A

Israel

Gorlova et al., 1997

Gladicosa pulchra (Keyserling, 1877)

Lycosa insopita

26+2S

X1X2

----

USA

Montgomery, 1905

Hippasa agelenoides (Simon, 1884)

----

28

X1X2

26A+X1X2A

India

Bole-Gowda, 1958

H. agelenoides (Simon, 1884)

----

24

X1X2

22A+X1X2A

India

Sharma et al., 1958

Hippasa holmerae Thorell, 1895

----

26

----

----

----

Bugayong et al., 1999

Hippasa madhuae Tikader & Malhotra, 1980

----

28

X1X2

26A+X1X2A

India

Parida et al., 1986

Hippasa olivacea (Thorell, 1887)

Hippasa oliracea

28

X1X2

26A+X1X2A

India

Parida & Sharma, 1987b

H. olivacea (Thorell, 1887)

----

28

X1X2

----

India

Parida & Sharma, 1987a

H. olivacea (Thorell, 1887)

----

28

X1X2

----

India

Sharma & Parida, 1987

Hippasa pisaurina Pocock, 1900

----

26

X1X2

----

India

Mittal, 1960

H. pisaurina Pocock, 1900

----

26

X1X2

24A+X1X2A

India

Mittal, 1963

H. pisaurina Pocock, 1900

----

28

X1X2

----

India

Srivastava & Shukla, 1986

Hippasa sp.

----

22

X1X2

----

India

Parida & Sharma, 1987a

Hippasa sp.

----

22

X1X2

----

India

Sharma & Parida, 1987

Hogna himalayensis (Gravely, 1924)

Lycosa himalayensis

28

X1X2

26A+X1X2A

India

Mittal, 1963

Hogna sternalis (Bertkau, 1880) ---- 19 X 18T+XT Brazil Araujo et al., 2015a

Hygrolycosa rubrofasciata (Ohlert, 1865)

----

20

X1X2

18A+X1X2A

Russia

Gorlov et al., 1995

Lycosa barnesi Gravely, 1924

----

27

X

----

India

Srivastava & Shukla, 1986

Lycosa bistriata Gravely, 1924

----

28

X1X2

26A+X1X2A

India

Bole-Gowda, 1958

Lycosa carmichaeli Gravely, 1924

----

28

X1X2

----

India

Mittal, 1961

L. carmichaeli Gravely, 1924

----

28

X1X2

26A+X1X2A

India

Mittal, 1963

L. carmichaeli Gravely, 1924

----

22

X1X2

----

India

Srivastava & Shukla, 1986

Lycosa chaperi Simon, 1885

----

22

X1X2

20A+X1X2A

India

Mittal, 1963

Lycosa coelestis L. Koch, 1878

----

26

X1X2

24A+X1X2A

Japan

Suzuki, 1954

Lycosa erythrognatha Lucas, 1836

Lycosa erythrognata

22

X1X2

20A+X1X2A

----

Diaz & Saez, 1966

L. erythrognatha Lucas, 1836

----

22

X1X2

20T+X1X2T

Argentina

Chemisquy et al., 2008

L. erythrognatha Lucas, 1836 ---- 22 X1X2 20T+X1X2T Brazil Araujo et al., 2015a

Lycosa madani Pocock, 1901

----

24

X1X2

22A+X1X2A

India

Mittal, 1963

Lycosa nigrotibialis Simon, 1884

----

28

X1X2

----

India

Mittal, 1961

L. nigrotibialis Simon, 1884

----

28

X1X2

26A+X1X2A

India

Mittal, 1963

L. nigrotibialis Simon, 1884

----

24

X1X2

----

India

Srivastava & Shukla, 1986

Lycosa nordenskjoldi Tullgren, 1905

Lycosa nordenskölii

19

X

18A+XA

----

Diaz & Saez, 1966

L. nordenskjoldi Tullgren, 1905 ---- 19 X 18T+XT Brazil Araujo et al., 2015a

Lycosa pampeana Holmberg, 1876

----

22

X1X2

20T+X1X2T

Argentina

Chemisquy et al., 2008

Lycosa phipsoni Pocock, 1899

----

----

X1X2

----

India

Mittal, 1961

Lycosa praegrandis C.L. Koch, 1836

Lycosa cf. nordmanni

22

X1X2

20A+X1X2A

Israel

Gorlova et al., 1997

Lycosa sericovittata Mello-Leitão, 1939 ---- 22 X1X2 20T+X1X2T Brazil Araujo et al., 2015a
L. sericovittata Mello-Leitão, 1939 ---- (26) ---- 26T Brazil Araujo et al., 2015a

Lycosa tarantula (Linnaeus, 1758)

Lycosa narborensis

18

----

----

Turkey

Akan et al., 2005

Lycosa thorelli (Keyserling, 1877)

----

22

X1X2

20T+X1X2T

Uruguay

Brum-Zorrilla & Postiglioni, 1980

L. thorelli (Keyserling, 1877)

----

22

X1X2

20T+X1X2T

Uruguay

Postiglioni & Brum-Zorrilla, 1981

Lycosa sp.

----

20-24?

----

----

France

Carnoy, 1885

Lycosa sp.

----

----

X

----

----

Chickering & Hard, 1935

Lycosa sp.

----

28

X1X2

26A+X1X2A

Japan

Suzuki, 1954

Lycosa sp.

----

28

X1X2

26A+X1X2A

India

Bole-Gowda, 1958

Lycosa sp.

----

28

X1X2

26A+X1X2A

India

Sharma et al., 1958

Lycosa sp.

----

28

X1X2

----

----

Sokolov, 1960

Lycosa sp.

----

28

X1X2

26A+X1X2A

India

Mittal, 1963

Lycosa sp.

----

22

X1X2

20T+X1X2T

Uruguay

Brum-Zorrilla & Postiglioni, 1980

Lycosa sp. (grupo malitiosa)

----

23

X

22T+XM

Uruguay

Postiglioni & Brum-Zorrilla, 1981

Lycosa sp. (grupo thorelli)

----

23

X1X2X3

20T+X1X2X3T

Uruguay

Postiglioni & Brum-Zorrilla, 1981

Lycosa sp.

----

22

X1X2

20T+X1X2T

Uruguay

Postiglioni & Brum-Zorrilla, 1981

Lycosa sp.

----

18

X1X2

----

India

Srivastava & Shukla, 1986

Lycosa sp.

----

28

X1X2

----

India

Srivastava & Shukla, 1986

Lycosa sp.

----

22

X1X2

20A+X1X2A

India

Parida & Sharma, 1987b

Lycosa sp.

----

22

X1X2

----

India

Parida & Sharma, 1987a

Lycosa sp.

----

28

X1X2

----

India

Parida & Sharma, 1987a

Lycosa sp.

----

22

X1X2

----

India

Sharma & Parida, 1987

Lycosa sp.

----

28

X1X2

----

India

Sharma & Parida, 1987

Lycosa sp.

----

21

X1X2Y

16T+2St+X1X2M/Sm+YT

Peru

Navia et al., 2006

Margonia himalayensis (Gravely, 1924)

Venonia himalayensis

28

X1X2

----

India

Mittal, 1961

M. himalayensis (Gravely, 1924)

Venonia himalayensis

28

X1X2

26A+X1X2A

India

Mittal, 1963

Ocyale kumari Dyal, 1935

----

22

X1X2

20A+X1X2A

India

Sharma et al., 1958

Pardosa agrestis (Westring, 1861)

----

28

X1X2

26A+X1X2A

Russia

Gorlov et al., 1995

Pardosa agricola (Thorell, 1856)

Lycosa fluviatilis

28

X1X2

26A+X1X2A

Finland

Hackman, 1948

Pardosa alacris (C.L. Koch, 1833)

----

28

X1X2

26A+X1X2A

Czech Republic

Kumbiçak et al., 2009

Pardosa amentata (Clerck, 1757)

Lycosa saccata

28

X1X2

26A+X1X2A

Finland

Hackman, 1948

P. amentata (Clerck, 1757)

Lycosa paludicola

28

X1X2

26A+X1X2A

Finland

Hackman, 1948

P. amentata (Clerck, 1757)

Lycosa pullata

28

X1X2

26A+X1X2A

Finland

Hackman, 1948

P. amentata (Clerck, 1757)

----

28

X1X2

----

----

Sokolov, 1960

Pardosa astrigera L. Koch, 1878

Lycosa T-insignita

28

X1X2

----

----

Suzuki, 1950a

P. astrigera L. Koch, 1878

Lycosa T-insignita

28

X1X2

26A+X1X2A

Japan

Suzuki, 1954

P. astrigera L. Koch, 1878

Lycosa cinereofusca

28

X1X2

26A+X1X2A

Japan

Suzuki, 1954

P. astrigera L. Koch, 1878

Pardosa T-insignita

28

X1X2

----

Japan

Matsumoto, 1977

P. astrigera L. Koch, 1878

----

28

X1X2

26T+X1X2T

Japan

Kageyama et al., 1978

P. astrigera L. Koch, 1878

----

28

X1X2

26T+X1X2T

China

Zhenling et al., 1996

Pardosa basiri (Dyal, 1935)

Lycosa basiri

22

X1X2

----

India

Mittal, 1960

P. basiri (Dyal, 1935)

Lycosa basiri

22

X1X2

20A+X1X2A

India

Mittal, 1963

Pardosa bifasciata (C.L. Koch, 1834) ---- 28 X1X2 26A+X1X2A Turkey Kumbiçak et al., 2011

Pardosa birmanica Simon, 1884

Lycosa birmanica

28

X1X2

26A+X1X2A

India

Bole-Gowda, 1958

P. birmanica Simon, 1884

----

28

X1X2

----

India

Datta & Chatterjee, 1983

P. birmanica Simon, 1884

Lycosa birmanica

28

X1X2

----

India

Srivastava & Shukla, 1986

P. birmanica Simon, 1884

----

28

X1X2

26A+X1X2A

India

Parida & Sharma, 1987b

P. birmanica Simon, 1884

----

28

X1X2

----

India

Parida & Sharma, 1987a

P. birmanica Simon, 1884

----

28

X1X2

----

India

Sharma & Parida, 1987

P. birmanica Simon, 1884

----

28

X1X2

26T+X1X2T

India

Datta & Chatterjee, 1989

Pardosa fletcheri (Gravely, 1924)

Lycosa fletcheri

28

X1X2

----

India

Srivastava & Shukla, 1986

Pardosa laura Karsch, 1879

----

28

X1X2

26T+X1X2T

Japan

Kageyama et al., 1978

Pardosa lugubris (Walckenaer, 1802)

----

28

X1X2

26A+X1X2A

Russia

Gorlov et al., 1995

Pardosa morosa (L. Koch, 1870)

----

26

X1X2

24A+X1X2A

Czech Republic/Slovakia

Král et al., 2011

P. morosa (L. Koch, 1870)

----

25?

X1X2?

24A+1M?

Czech Republic

Král et al., 2011

Pardosa palustris (Linnaeus, 1758)

Lycosa monticola

28

X1X2

26A+X1X2A

Finland

Hackman, 1948

P. palustris (Linnaeus, 1758)

Lycosa tarsalis

28

X1X2

24A+2M+X1X2A

Finland

Hackman, 1948

P. palustris (Linnaeus, 1758)

----

28

X1X2

26A+X1X2A

Russia

Gorlov et al., 1995

Pardosa plumipes (Thorell, 1875)

----

28

X1X2

26A+X1X2A

Russia

Gorlov et al., 1995

Pardosa pseudoannulata (Bösenberg & Strand, 1906)

Lycosa pseudoannulatus

28

X1X2

26A+X1X2A

Japan

Suzuki, 1954

P. pseudoannulata (Bösenberg & Strand, 1906)

Lycosa annandalei

28

X1X2

26A+X1X2A

India

Bole-Gowda, 1958

P. pseudoannulata (Bösenberg & Strand, 1906)

Lycosa pseudoannulata

28

X1X2

26T+X1X2T

Japan

Kageyama et al., 1978

P. pseudoannulata (Bösenberg & Strand, 1906)

Lycosa annandalei

28

X1X2

----

India

Srivastava & Shukla, 1986

Pardosa saltans Töpfer-Hofmann, 2000

----

28

X1X2

26A+X1X2A

Czech Republic

Kumbiçak et al., 2009

Pardosa sumatrana (Thorell, 1890)

Lycosa sumatrana

24

X1X2

22A+X1X2A

India

Sharma, 1961

P. sumatrana (Thorell, 1890)

Lycosa sumatrana

28

X1X2

----

India

Srivastava & Shukla, 1986

Pardosa wagleri (Hahn, 1822) ---- 28 X1X2 26A+X1X2A ---- Kořínkova & Král, 2013

Pardosa sp.

----

----

X

----

----

Chickering & Hard, 1935

Pardosa sp.

----

28

X1X2

26T+X1X2T

India

Sharma & Gupta, 1956

Pardosa sp.

----

28

X1X2

26A+X1X2A

India

Bole-Gowda, 1958

Pardosa sp.

----

28

X1X2

----

India

Mittal, 1960

Pardosa sp.

----

28

X1X2

26A+X1X2A

India

Mittal, 1963

Pirata piraticus (Clerck, 1757)

----

26

X1X2

24A+X1X2A

Finland

Hackman, 1948

P. piraticus (Clerck, 1757)

----

26

X1X2

24T+X1X2T

----

Yu et al., 1998a

Pirata subpiraticus (Bösenberg & Strand, 1906)

----

26

X1X2

24T+X1X2T

Japan

Kageyama et al., 1978

P. subpiraticus (Bösenberg & Strand, 1906)

----

26

X1X2

24T+X1X2T

----

Yu et al., 1998b

Pirata sp.

----

26

X1X2

----

----

Sokolov, 1960

Piratula latitans (Blackwall, 1841)

Pirata latitans

24

X1X2

22A+X1X2A

India

Mittal, 1963

Piratula piratoides (Bösenberg & Strand, 1906)

Pirata piratoides

26

X1X2

24T+X1X2T

----

Zhenling et al., 1997a

Piratula procurva (Bösenberg & Strand, 1906)

Pirata procurvus

26

X1X2

24T+X1X2T

Japan

Kageyama et al., 1978

Piratula uliginosa (Thorell, 1856)

Pirata uliginosus

24

X1X2

22A+X1X2A

Finland

Hackman, 1948

Rabidosa rabida (Walckenaer, 1837)

Lycosa rabida

28

X1X2

26T+X1X2T

USA

Wise, 1983

R. rabida (Walckenaer, 1837)

Lycosa rabida

28

X1X2

----

USA

Tugmon et al., 1990

R. rabida (Walckenaer, 1837)

Lycosa rabida

28

X1X2

26T+X1X2T

USA

Wise & Taylor, 1995

Schizocosa bilineata (Emerton, 1885)

----

----

X

----

----

Chickering & Hard, 1935

Schizocosa communis (Emerton, 1885)

Lycosa communis

22

X1X2

----

----

Painter, 1914

Schizocosa crassipes (Walckenaer, 1837)

----

22

X1X2

----

USA

Hard, 1939

Schizocosa malitiosa (Tullgren, 1905)

Lycosa malitiosa

22

X1X2

20T+X1X2T

Uruguay

Brum-Zorrilla & Cazenave, 1974

S. malitiosa (Tullgren, 1905)

Lycosa malitiosa

22

X1X2

----

Uruguay

Benavente & Wettstein, 1977

S. malitiosa (Tullgren, 1905)

Lycosa malitiosa

22

X1X2

20T+X1X2T

Uruguay

Brum-Zorrilla & Postiglioni, 1980

S. malitiosa (Tullgren, 1905)

----

22

X1X2

20T+X1X2T

Argentina

Chemisquy et al., 2008

Schizocosa ocreata (Hentz, 1844)

----

22

X1X2

----

USA

Stratton, 1997

Schizocosa rovneri Uetz & Dondale, 1979

----

22

X1X2

----

USA

Stratton, 1997

Schizocosa stridulans Stratton, 1984

----

22

X1X2

----

USA

Stratton, 1997

Schizocosa sp.

----

28

X1X2

----

India

Mittal, 1960

Schizocosa sp.

----

28

X1X2

26A+X1X2A

India

Mittal, 1963

Tigrosa georgicola (Walckenaer, 1837)

Lycosa georgicola

28

X1X2

24T+2A+X1X2T

USA

Wise, 1983

Tigrosa helluo (Walckenaer, 1837)

Lycosa helluo

28

X1X2

26T+X1X2T

USA

Wise & Taylor, 1995

Trochosa punctipes (Gravely, 1924)

Lycosa punctipes

28

X1X2

26A+X1X2A

India

Sharma, 1961

Trochosa ruricola (De Geer, 1778)

----

26

X1X2

24A+X1X2A

Finland

Hackman, 1948

T. ruricola (De Geer, 1778) Trachosa ruricula 20 X1X2 ---- Turkey Taşdemir et al., 2012

Trochosa spinipalpis (F.O.Pickard-Cambridge, 1895)

----

26

X1X2

24A+X1X2A

Finland

Hackman, 1948

T. spinipalpis (F.O.Pickard-Cambridge, 1895)

----

26

X1X2

24A-T+X1X2A-T

----

Zhenling et al., 1997b

Venonia sp.

----

26

X1X2

----

India

Mittal, 1961

Venonia sp.

----

26

X1X2

----

India

Mittal, 1961

Venonia sp.

----

26

X1X2

24A+X1X2A

India

Mittal, 1963

Wadicosa fidelis (O.Pickard-Cambridge, 1872) ---- 28 X1X2 26A+X1X2A Canary Island, Israel Forman et al., 2013

Wadicosa quadrifera (Gravely, 1924)

Lycosa quadrifer

27

X

----

India

Srivastava & Shukla, 1986

Xerolycosa miniata (C.L. Koch, 1834)

----

22

X1X2

20A+X1X2A

Finland

Hackman, 1948

X. miniata (C.L. Koch, 1834)

----

22

X1X2

20A+X1X2A

Russia

Gorlov et al., 1995

X. miniata (C.L. Koch, 1834)

----

22

X1X2

20A+X1X2A

Czech Republic

Dolejš et al., 2011

Xerolycosa nemoralis (Westring, 1861)

----

26

X1X2

24A+X1X2A

Finland

Hackman, 1948

X. nemoralis (Westring, 1861)

----

22

X1X2

20A+X1X2A

Russia

Gorlov et al., 1995

X. nemoralis (Westring, 1861)

----

22

X1X2

20A+X1X2A

Czech Republic

Dolejš et al., 2011

 


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