The scorpion cytogenetic database

Current version: 9.5 (Jan 13, 2021)

Marielle Cristina Schneider, Viviane Fagundes Mattos and Doralice Maria Cella


Introduction of Database Families - Cytogenetics of spiders Bibliography Arthropoda cytogenetics group Spider database Pseudoscorpion database Harvestmen database Chromosomal analyses

Bibliography

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ADILARDI, R. S.; AFFILASTRO, A. O.; MARTÍ, D. A.; MOLA, L. M. Cytogenetic analysis on geographically distant parthenogenetic populations of Tityus trivitattus Kraepelin, 1898 (Scorpiones, Buthidae): karyotype, constitutive heterochromatin and rDNA localization. Comparative Cytogenetics, v. 8, p. 81-92, 2014.

 

ADILARDI, R. S.; OJANGUREN-AFFILASTRO A. A.; MATTONI, C. I.; MOLA, L. M. Male and female meiosis in the mountain scorpion Zabius fuscus (Scorpiones, Buthidae): heterochromatin, rDNA and TTAGG telomeric repeats. Genetica. v. 143, p.393-401, 2015.

 

ADILARDI, R. S.; OJANGUREN-AFFILASTRO, A. A.; MOLA, L. M. Sex-linked chromosome heterozigosity in males of Tityus confluens (Buthidae): a clue about the presence of sex chromosomes in scorpions. PLoS ONE, v. 11, n. 10, p. 1-18, 2016.

 

ADILARDI, R.S.; OJANGUREN-AFFILASTRO, A.A.; MARTÍ, D.A.; MOLA, L.M. Chromosome puzzle in the southernmost populations of the medically important scorpion Tityus bahiensis (Perty 1833) (Buthidae), a polymorphic species with striking structural rearrangements. Zoologischer Anzeiger. v. 288, p. 139-150, 2020.

 

ALMEIDA, B.R.R.; MILHOMEM-PAIXÃO, S.S.R.; NORONHA, R.C.R.; NAGAMACHI, C.Y.; COSTA, M.J.R.; PARDAL, P.P.O.; COELHO, J.S.; PIECZARKA, J.C. Karyotype diversity and chromosomal organization of repetitive DNA in Tityus obscurus (Scorpiones, Buthidae). BMC Genetics n. 18, 2017.

 

Almeida, B.R.R.; Noronha, R.C.R.; Costa, M.R.J.; Nagamachi, C.Y.; Pieczarka, J.C. Meiosis in scorpion Tityus silvestris: new insigths into achiasmatic chromosome. Biology Open. v. 8, p. bio040352, 2019.

 

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FERREIRA A. Contribution to the knowledge of cytology of two species of Brazilian scorpions: Opisthacantus manauarensis, Ferreira, 1967 (Scorpiones, Scorpionidae) and Bothriurus asper araguaie (Scorpiones, Bothriuridae). Anais da Academia Brasileira de Ciências. v. 40, p. 97-99, 1968.

 

GUÉNIN, H. A. Contribution à la connaissance cytologique des scorpions: les chromosomes de Pandinus imperator Koch. Revue Suisse de Zoologie, v. 64, p. 349-353, 1957.

 

GUÉNIN, H. A. Contribution a la connaissance cytologique des scorpions: les chromosomes de Buthus occitanus Amor. (I). Vie et Milieu. v. 12, p. 89-96, 1961.

 

GUPTA, A. K. D. and SARKER, S. K. A study of the meiotic chromosomes of the scorpion Buthus tamulus Fabr. Current Science. v. 2, p. 54-55, 1965.

 

KARATAŞ, A., UÇAK, M., KARATAŞ, A. and EROĞLU, O. Records of chromosomal data of some scorpions (Arachnida: Scorpiones) from Turkey. Hacettepe Journal of Biology and Chemistry, v. 47, p. 1-6, 2019.

 

KOÇ, H., YAGMUR, E.A. and ŠŤÁHLAVSKÝ, F. Karyotypes of Southeastern Turkish Scorpions Hottentotta saulcyi and Buthacus macrocentrus (Scorpiones: Buthidae). European Journal of Biology, v. 78(2), p. 1-6, 2019.

 

KOVAŘÍK, F. and ŠŤÁHLAVSKÝ, F. Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius, v. 315, p. 1-37, 2020.

 

KOVAŘÍK, F.; STAHLAVSKÝ, F.; KORÍNKOVÁ, T.; KRÁL, J. and ENDE, T. Tityus ythieri Lourenço, 2007 is a synonym of Tityus magnimanus Pocock, 1897 (Scorpiones: Buthidae): a combined approach using morphology, hybridization experiments, chromosomes, and mitochondrial DNA. Euscorpius, v. 77, p. 1-12, 2009.

 

KOVAŘÍK, F.; PLISKOVA, J. and STAHLAVSKY, F. Euscorpiops neradi sp. n. from  Thailand (Scorpiones: Euscorpiidae: Scorpiopinae). Euscorpius,  n. 173, p. 01-19, 2013.

 

KOVAŘÍK, F.; KRÁL, J.; KOŘÍNKOVÁ, T. and LERMA, A.C.R. Chaerilus hofereki sp. n. from Vietnam (Scorpiones: Chaerilidae). Euscorpius, n. 189, p.1–11 2014.

 

KOVAŘÍK, F.; LOWE, G.; SEITER, M.; PLISKOVÁ, J.; STAHLAVSKY, F. Scorpions of Ethiopia (Arachnida: Scorpiones). Part II. Genus Babycurus Karsh, 1886 (Buthidae), with description of two new species. Euscorpius. v. 196, p. 1-31, 2015a.

 

KOVAŘÍK, F.; KOSULIC, O.; STAHLAVSKY, F.; PLISKOVÁ, J.; DONGKHAMFU, W.; WONGPROM, P. Two new species of Euscorpiops Vachon, 1980 from Thailand and Myanmar (Scorpiones: Euscorpiidae: Scorpiopinae). Annales Zoologici (Warszawa). v.65, p. 109-122, 2015b.

 

KOVAŘÍK, F.; LOWE, G.; HOFEREK, D.; FORMAN, M. and KRÁL, J. Two new Chaerilus from Vietnam (Scorpiones, Chaerilidae), with observations of growth and maturation of Chaerilus granulatus sp. n. and C. hofereki Kovařík et al., 2014. Euscorpius, n. 213, p. 1-21, 2015c.

 

KOVAŘÍK, F., SOLEGLAD, M.E., LOWE, G., PLÍŠKOVÁ, J. and ŠŤÁHLAVSKÝ, F. Observations on Growth and Maturation of a Male Alloscorpiops wongpromi (Scorpiones: Euscorpiidae: Scorpiopinae: Scorpiopini). Euscorpius, v. 205, p. 1-17, 2015d.

 

KOVAŘÍK, F.; LOWE, G. and ŠŤÁHLAVSKÝ, F. Review of northwestern African Buthacus, with description of Buthacus stockmanni sp. n. from Morocco and western Sahara (Scorpiones, Buthidae). Euscorpius. n. 236, p. 1-18, 2016a.

 

Kovarík, F.; Lowe, G.; Ranawana, K.B.; Hoferek, D.; Jayarathne, V.A.S.; Plisková, J.; Stahlavský, F. Scorpions of Sri Lanka (Arachnida, Scorpiones, Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972, stat. n. Euscorpius n. 220, p.1-133, 2016b.

 

Kovarík, F.; Lowe, G.; Plisková, J.; Stahlavský F. Scorpions of horn of Africa (Arachnida: Scorpiones). Part VI. Compsobuthus Vachon, 1949 (Buthidae), with a description of C. eritreaensis sp. n. Euscorpius n. 226, p. 1-21, 2016c.

 

KOVAŘÍK,F.; LOWE, G.; PLÍŠKOVÁ, J. and ŠŤÁHLAVSKÝ, F. Scorpions of the horn of Africa (Arachnida:Scorpiones). Part VII. Parabuthus Pocock, 1980 (Buthidae), with description of P. hamar sp. n. and P. kajibu sp. n. from Ethiopia. Euscorpius. n.228, p. 1-58, 2016d.

 

KOVAŘÍK, F; LOWE, G. and ŠŤÁHLAVSKÝ, F. Scorpions of the horn of Africa (Arachnida: Scorpiones). Part. IX. Lanzatus, Orthochirus, and Somalicharmus (Buthidae), with description of Lanzatus somalilandus sp. n. and Orthochirus afar sp. n. Euscorpius. n. 232, p. 1-38, 2016e.

 

Kovarík, F.; Lowe, G.; Hoferek, D.; Plisková, J.; Stahlavský, F. Scorpions of Ethiopia. Part IV. Genus Uroplects Peters, 1861 (Scorpiones: Buthidae). Euscorpius n. 217, p.1-14, 2016f.

 

KOVARÍK, F.; LOW, G.; MAZUCH, T.; PLISKOVÁ, J.; STAHLAVSKÝ F. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XI. Pandinurus kmoniceki sp. n. (Scorpionidae) from Somaliland. Euscorpius n. 243, p. 1-14, 2017a.

 

Kovařík, F., LOWE, G., MAZUCH, T., AWALE, A.I., Štundlová, J., Šťáhlavský, F. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XII. Pandinurus hangarale sp. n. (Scorpionidae) from Somaliland and a review of type locality and true distribution of Pandinurus smithi (Pocock, 1897). Euscorpius. n. 253, p.1-18, 2017b.

 

KOVAŘÍK, F.; LOWE, G.; ŠŤÁHLAVSKÝ, F. Review of the genus Babycurus Karsch, 1886 (Arachnida, Scorpiones, Buthidae), with descriptions of Barbaracurus gen. n. and two new species from Oman and Yemen. Euscorpius. n. 267, p. 1-41, 2018a.

 

KOVAŘÍK, F.; LOWE, G.; ŠŤÁHLAVSKÝ, F. Three new Chaerilus from Malaysia (Tioman Island) and Thailand (Scorpiones: Chaerilidae), with a review of C. cimrmani, C. sejnai, and C. tichyi. Euscorpius. n. 268, p. 1-27, 2018b.

 

KOVAŘÍK, F., LOWE, G., JUST, P., AWALE, A. I., ELMI, H.S.A and ŠŤÁHLAVSKÝ, F. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XV. Review of the genus Gint Kovařík et al, 2013, with description of three new species from Somaliland (Scorpiones, Buthidae). Euscorpius, v. 259,  p. 1-41, 2018c.

 

Kovařík, F.; Šťáhlavský, F. Revision of the genus Reddyanus from Southeast Asia, with description of five new species from Cambodia, Malaysia, Thailand and Vietnam (Scorpiones: Buthidae). Euscorpius - Occasional Publications in Scorpiology, n. 295, 2019.

 

Kovařík, F.; Lowe, G.; Šťáhlavský, F.; Hurre, A.A. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XX. Barbaracurus feti sp. n. from Somaliland (Buthidae). Euscorpius - Occasional Publications in Scorpiology, n. 280, 2019a.

 

Kovařík, F.; Ranawana, K.B.; Jayarathne, V.A.S.; Hoferek, D.; Šťáhlavský, F. Scorpions of Sri Lanka (Arachnida: Scorpiones). Part III. Heterometrus yaleensis sp. n. (Scorpionidae). Euscorpius - Occasional Publications in Scorpiology, n. 283, 2019b.

 

Kovařík, F.; Štundlová, J.; Fet, V.; Šťáhlavský, F. Seven new Alpine species of the genus Alpiscorpius Gantenbein et al., 1999, stat. n. (Scorpiones: Euscorpiidae). Euscorpius - Occasional Publications in Scorpiology, n. 287, 2019c.

 

KOVAŘÍK, F., LOWE, G., ELMI, H.S.A. and ŠŤÁHLAVSKÝ, F. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XXI. Parabuthus (Buthidae) (Part II), with description of five new species from Somaliland and Ethiopia. Euscorpius, v. 290, p. 1-6, 2019d.

 

KOVAŘÍK, F., LOWE, G., STOCKMANN, M., and ŠŤÁHLAVSKÝ, F. Notes on Compsobuthus: redescription of C. arabicus Levy et al., 1973 from Arabia, and description of two new species from North Africa (Scorpiones: Buthidae). Euscorpius, n. 298, 2020a.

 

KOVAŘÍK, F., ŠŤÁHLAVSKÝ, F. and ELMI, H.S.A. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XXIII. Buthus (Buthidae), with description of two new species. Euscorpius, n. 307, 2020b.

 

Kovařík, F.; Lowe, G.; Šťáhlavský, F. Reddyanus justi sp. n. from Laos. Euscorpius. v. 321, p. 1-11, 2020c.

 

KOVAŘÍK, F.; LOWE, G.; STOCKMAN, M.; ŠŤÁHLAVSKÝ, F. Two new Chaerilus from Thailand and Laos (Scorpiones: Chaerilidae). Euscorpius. v. 324, p. 1–20, 2020d.

 

Kovařík, F.; Lowe, G.; Stockmann, M; Šťáhlavský, F. Revision of genus-group taxa in the family Scorpiopidae Kraepelin, 1905, with description of 15 new species (Arachnida: Scorpiones). Euscorpius. v. 325, p. 1–140, 2020e.

 

Kovařík, F.; Lowe, G.; Byronová, M.; Šťáhlavský, F. Euscorpius thracicus sp. n. (Scorpiones: Euscorpiidae) from Bulgaria. Euscorpius. v. 326, p. 1–17, 2020f.

 

LOWE, G. KOVARIK, F. STOCKMANN, M. STAHLAVSKY, F.  Review of Microbuthus with description of M. satyrus sp. n. (Scorpiones, Buthidae) from Oman and Yemen. Euscorpius,  n. 263, p. 1-22, 2018.

 

MAKINO, S. A review of the chromosome numbers in animals. Tokyo: Hokurykan. 1956.

 

MATTOS, V. F.; CELLA, D. M.; CARVALHO, L. S.; CANDIDO, D. M.; SCHENEIDER, M. C. High chromosome variability and the presence of multivalent association in buthid scorpions. Chromosome Research, v. 21, p. 121-136, 2013.

 

MATTOS, V. F. CARVALHO, L. S. CARVALHO, M. A. SCHNEIDER, M. C. Insigths into the origin of the high variability of multivalent meiotic associations in holocentric chromosomes of Tityus (Archaeotityus) scorpions. PloS One, v. 13, p. e0192070, 2018.

 

MOUSTAFA, M. A.; ALAA, A. M.; SARHAN, M. H. and YASEEN, A. E. Chromosomal studies on four Egyptian scorpion species of genus Androctonus (Family: Buthidae). Cytologia, v. 70, p. 161-165, 2005.

 

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PIZA, S. T. Meiosis in the male of the Brazilian scorpion Tityus bahiensis. Revista de Agricultura. v. 18, p. 249-276, 1943a.

 

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PIZA, S. T. Notas sobre cromossômios de alguns escorpiões brasileiros. Anais da Escola Superior de Agricultura “Luiz de Queiroz”, v. 62, p. 169-176, 1947a.

 

PIZA, S. T. Observações cromossômicas em escorpiões brasileiros. Ciência e Cultura. v. 2, p. 202-206, 1950b.

 

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PIZA, S. T. Primeiras observações sôbre os cromossômios do Tityus metuendus Pocock. Scientia Genetica. v. 4, p. 162-167, 1952.

 

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PIZA, S. T. Variações cromossômicas do Tityus bahiensis de Ribeirão Preto. Ciência e Cultura. v. 2, p. 57-59, 1950a.

 

PIZA, S. T. Variações cromossômicas do Tityus bahiensis de São Joaquim. Revista de Agricultura de São Paulo. v. 24, p. 187-194, 1848c.

 

PLÍŠKOVÁ, J.; KOVAŘÍK, F.; KOŠULIČ, O. and ŠŤÁHLAVSKÝ, F. Description of a new species of Heterometrus Ehrenberg, 1828 (Scorpiones: Scorpionidae) from Thailand with remarks about the utilization of cytogenetic data in taxonomy of the genus. Annales Zoologici. v. 66, n. 3, p. 467-476, 2016.

 

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QUMSIYEH, M. B.; AMR, Z. S.; SROUR, K. T. A.; AL-FAWAGHRA, N. Karyotype for Nebo hierichonticus (Simon 1872) from the Palestinian Territories (Scorpiones: Scorpiopidae). Cytologia. v.79, 277-280, 2014.

 

RAJASEKARASETTY, M. R.; ASWATHANARAYANA, N. V. and KUMARASWAMY, K. R. Chromosome biology of the scorpion Palamnaeus sp. Proceedings of the Indian Academy of Sciences. v. 88, p. 187-192, 1979.

 

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RODRÍGUEZ-GIL, S. G.; OJANGUREN-AFFILASTRO, A. A.; BARRAL, L. M.; SCIOSCIA, C. L. and MOLA, L. M. Cytogenetic of three species of the genus Brachistosternus from Argentina (Scorpiones: Bothriuridae). The Journal of Arachnology. v. 37, p. 331-337, 2009.

 

SADÍLEK, D.; NGUYEN, P.; KOÇ, H.; KOVARÍK, F.; YAGMUR, E. A.; STAHLAVSKÝ, F. Molecular cytogenetics of Androctonus Scorpions: an oasis of calm in the turbulent karyotype evolution of the diverse family Buthidae. Biological Journal of the Linnean Society. v. 115, p. 69-76, 2015.

 

SATO, I. Studies on the cytoplasmic phenomena in the spermatogenesis of the Oriental scorpion, Buthus martensii, with special reference to the structure of the chondriosome ring and the dictyokinesis. Journal of Science of the Hirosima University. v. 8, p. 1-116, 1940.

 

SCHNEIDER, M. C. and CELLA, D. M. Karyotype conservation in 2 populations of the parthenogenetic scorpion Tityus serrulatus (Buthidae): rDNA and its associated heterochromatin are concentrated on only one chromosome. Journal of Heredity. v. 101, p. 491-496, 2010.

 

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SRIVASTAVA, M. D. L. and AGRAWAL, U. Absence of chiasmata and formation of a complex chromosomal body in the spermatogenesis of the scorpion Palamnaeus longimanus. Caryologia. v. 14, p. 63-77. 1961.

 

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Šťáhlavský, F.; Nguyen; P.; Sadílek, D.; Štundlová, J.; Just, P.; Haddad, C.R.; Koc, H.; Ranawana, K.B.; Stockmann, M.; Yagmur, E.A.; Kovařík, F. Evolutionary dynamics of rDNA clusters on chromosomes of buthid scorpions (Chelicerata: Arachnida). Biological Journal of the Linnean Society. v. 131, p. 547-565, 2020.

 

Štundlová, J.; Šmíd, J.; Nguyen, P.; Šťáhlavský, F. Cryptic diversity and dynamic chromosome evolution in Alpine scorpions (Euscorpiidae: Euscorpius). Molecular Phylogenetics and Evolution. v. 134, p. 152-163, 2019.

 

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UBINSKI, C. V.; CARVALHO, L. S.; SCHNEIDER, M. C. Mechanisms of karyotype evolution in the brazilian scorpions of the subfamily Centruroidinae (Buthidae). Genetica. v.146, n. 6, p. 475-486, 2018.

 

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Introduction of Database Families - Cytogenetics of spiders Bibliography Arthropoda cytogenetics group Spider database Pseudoscorpion database Harvestmen database Chromosomal analyses